بررسی ویژگیهای مورفولوژیکی و کمی و کیفی توده‌های راش ایران از دیدگاه حفاظت ژن

نوع مقاله : علمی- پژوهشی

نویسندگان

1 اعضاء هیأت علمی مؤسسه تحقیقات جنگلها و مراتع‏

2 اعضاء هأت علمی مؤسسه تحقیقات جنگلها و مراتع‏

چکیده

گونه راش یکی از فراوانترین و از نظر اقتصادی مهمترین درختان جنگلهای شمال ایران است. تعیین ساختار هر توده و ترکیب آن با خصوصیات جنگل‌شناسی که شامل آمیختگی، سن، ساختار عمودی یا اشکوب‌بندی، ساختار افقی، وجود یا عدم وجود روزنه، تاج پوشش، انبوهی، وجود و یا عدم وجود زادآوری یا زیراشکوب درختی و درختچه‌ای است می‌تواند ما را در دستیابی به یک مدیریت مناسب و فراگیر، یاری نماید.
در این پژوهش خصوصیات مورفولوژیکی و ویژگیهای ژنتیکی در 14 قطعه نمونه نیم هکتاری در راشستانهای شمال ایران مورد بررسی قرار گرفته است. هم چنین سعی شده است نقش عوامل انسانی و مدیریت راشستانهای مورد نظر (شیوه‌های جنگل‌شناسی بکار رفته) بر این خصوصیات مطالعه شود.
 
تنوع ژنتیکی بالای راش در راشستانهای ایران (هتروزیگوزیتی مورد انتظار= 191/0) حاکی از توان سازگاری این درخت در جنگلهای خزری می‌باشد. از آنجایی که رابطه مثبتی بین تنوع ژنتیکی و درصد پایه‌های چند شاخه و غیرمستقیم (بد فرم) وجود دارد، بنابراین از لحاظ نظری انتخاب و حفظ درختان خوش فرم برای زادآوری در توده‌ها که در اثر فعالیتهای مدیریت تحت شیوه تدریجی پناهی اعمال می‌گردد می‌بایست با احتیاط با آن برخورد شود. زیرا انتخاب برای حفظ برخی ژنوتیپها از مهمترین فرایندهایی است که در طی عملیات مدیریت جنگل باعث تغییر در گوناگونی ژنتیکی درختان می‌گردد. در طی این فرایند میزان برخی ژنها به شدت کاهش می‌یابد و بنابراین، باعث کاهش گوناگونی ژنتیکی می‌شود که می‌تواند روی سازگاری آینده تأثیر بگذارد. بنابراین یک شیوه جنگل‌شناسی مناسب می‌بایست ضمن حفاظت توده‌ها، همزمان با برداشت و استفاده از محصولات و خدمات جنگل در حال حاضر، کوششی برای تأمین محصولات، کالاها و سایر خدمات در آینده نیز باشد. به این ترتیب میزان و چگونگی استفاده از منابع موجود در حدی خواهد بود تا تأمین مناسب و مستمر آنها در آینده تضمین گردد. همان گونه که مطالعات اخیر نشان داده‌اند در میان شیوه‌های جنگل‌شناسی دو شیوه تک گزینی و شیوه گروه‌گزینی با ساختار توده‌های طبیعی راش شمال ایران همخوانی بیشتری دارد. در این پژوهش نیز توان سازگاری بالای برخی مناطق (نکا در ارتفاع 900 متر و خیرود در ارتفاع 600 متر از سطح دریا با 14 آلل نادر و مختص به محل) در مقایسه با سازگاری محدود برخی دیگر (مثل خیرود در ارتفاع 1200 متر و گرگان در ارتفاع 2000 متر از سطح دریا با 2 آلل نادر و مختص به محل) تأکیدی بر حفاظت راشستانهای شمال ایران و انتخاب شیوه جنگل‌شناسی مناسب با هدف تولید مستمر و حفاظت از تنوع ژنتیکی توده‌های راش می‌باشد.
Determination of beech (Fagus orientalis Lipsky)
seeding cycle in Ziarat Forest
 
S. Z. Mirkazemi1 and A. R. Mirbadin2
 
Abstract
The trail was conducted at Ziarat Forest (Longitude: 54º 25´ East, Latitude: 36º 43´ North, altitude: 650-2350. asl) of Caspian region of Iran in order to identify the seedling cycle of F. orientalis.
Twenty seven dominant beech trees were selected for quantitative and qualitative measurment. After soil and ground cover sampling and tree measurment, the trees were cut down and a disc sample was taken from each tree's stump. The discs were analyzed to study the annual rings. The results showed that:
1-Fagus orientalis has two seeding types, heavy and light.
2- The minimum age of seeding is about 49 year when is often associated with light seeding.
3- The heavy seeding begin at age of 57 year.
4- At a same site, different seeding cycles were observed during the living period.
5- The light and the heavy seeding repeat at 2-6 and 3-27 year period, respectively.
6- Natural regeneration depends on the light seeding cycles due to their frequent occurance.
7- There was a relationship between the averag decerese of annual ring growth rate and seeding types as follows:
a. If decrease of annual ring growth rate is more than 1mm, there will be a heavy seeding cycle.
b. If decrease of annual ring growth rate is between 1mm and 0.31mm, then there will be alight seeding cycle.
C. If decrease of annual ring growth rate is less than 0.31mm, there will not be a seeding cycle.
 
Key words: Forest, beech, Fagus orientalis, seeding cycle, growth, age.


Effects of eucalypt plantation on understory plant cover and some soil parameters in comparison to next natural forest in Astara of Iran
 
F.Moraghebi1, S. A. A. Korori2, B. Khanjani-Shiraz3, M. Teimouri2
 and A. Hemmati3
 
Abstract
About 30 years ago, a eucalypt species trial was conducted in Caspian coastal forest of Iran at Astara, after clear felling the natural QuercoCarpinetum forest type. The aim of the study was investigation the effect eucalypt plantation on understory plant cover component and soil properties .For this purpose, the understory species and their frequency was determined in both eucalyptus plantation and next natural forest. The Sorenson method was used to study the heterogeneity homogeneity of plant communities .Furthermore, the soil Na+, K+, organic matter and electrical conductivity (EC) were measured. Results showed that there was only 54% homogeneity between the understory plant cover of two sites according to Sorenson index that indicated the significant effects of Eucalypt plantation on kind and frequency of species. The amounts of Na+, K+, organic matter and EC was less in eucalypt plantation site  than the natural forest site.
The results showed that there was only 54% homogeneity between the understory plant cover of the two sites according to Sorenson Index and significant effect of eucalypt plantation on flora. The amount of Na+, K+, EC and organic matter in eucalypt plantation site was less than the natural forest site. It was concluded that eucalypt plantation on natural forest sites should be limited due to its significant effects on sol and flora.
 
Key words: Forestation, Eucalyptus, natural forest, Plant understory.


Effects of using Aquasorb Polymer on irrigation of Pinus eldarica, Olea europea and Atriplex canescens
 
A. Pourmeidani1 and H. Khakdaman1
 
Abstract
This experiment was carried out in 2001 in order to determine the effect of aquasorb polymer on irrigation of three plant species in Hossein Abad Station of Sand Stabilization of Qom province.
The Factorial statistical method under Randomized compete blocks design whit three replicates was used. There were two Factors: species (Atriplex canescens, Pinus eldarica Medw. and Olea europea L.) and Aquasorb polymer (0.0%, 2.0%, 4.0% with irrigation interval three times were than normal interval) and control irrigation with normal interval). The variables were: Soil water content (%) before and after irrigation (at depths of 25 and 50 cm) and survival % one, three and six months after plating. The ANOVA and Duncan tests were used to analyze the data.
The results showed that there were significant differences (p<0.01) in respect to survival% between the planted species. The species Atriplex canescens p., Olea europea and Pinus eldarica obtained the greatest survival percentage, respectively. The effects of Aquasorb polymer treatments on the measured variables (except soil water content ratio of after irrigation to before irrigation at 50 cm depth) were significant (p<0.01). There was a not significant differences between the levels 2.0 and 4.0% of Aquasorb polymer on one hand and between these levels and control irrigation regime on another hand. It is suggested to apply Aquasorb at level of 2.0 instead of normal irrigation for planting three species at similar climate and soil conditions as the studied site.
 
Key words: Aquasorb; irrigation, polymer, Pinus, Olea, Atriplex


Distribution of different species of Pistachio in Markazi Province of Iran
 
H. Zahedipour1, M. Fattahi2, H. R. Mirdavoodi Akhavan1,
Gh. R. Godarzi1 and Z. Azdoo3
 
Abstract
Considering appropriate adaptability of different wild species of Pistacia in different arid and Semi-arid environments of the country, as well as their high value from point of view of Resin and other chemical production; oil; and pharmacological use of their seeds, studying their habitats in different areas of Markazi Province, (as a sub-project of the national project under the title of “The study of Pistacia distribution and related influencing factors in Iran) seems to be important. In this project the basic maps including topography, soil, climate, geology, and landuse of the area (1:50000 scale) were purchased and digitized. Then the areas which were covered by the wild species were visited, recorded by GPS and digitized. In order to study the quantitative and qualitative characteristics of the species, five sample plots (1600m2) The tree parameters were: density, total height, stem diameter, crown height, crown diameter, type of regeneration, stem and crown health, number of male and female Pistachios, amount of seed production and seed quality. After establishing the DEM of the areas, aspect and slope maps were also produced using dfdx and dfdy filters of IlWIS Academic Program Package.
On the basis of this investigation the following results were obtained.
-Twenty polygons of wild Pistacia species of (areas of 2216 ha) were recognized in Markazi Province.
- The species - environment relationships including: slop, aspect, climate, landuse and soil were determined, using maps integrating and data overlying methods.
 
Key words: Pistachia atlantica, Pistachia Khinjuk, Amygdalus, tree parameter.


Survival and growth of 10 broadleaved and needleleaved tree species in Fars Province
 
M. R. Negahdar - Saber1, M. Hamzehpour1 and L. Joukar1
 
Abstract
Importance of wood as a basic material in industry, along its broad uses and diverse by-products leads us to change our direction from exploitation of natural forests towards artificial forest plantation. This is done mainly by selecting suitable and fast-growing species. Understanding the adaptation and the growth potential of forest trees in different environments is one of the earliest steps in approaching this great goal.
In this study, growth and survival trend of ten forest tree species including Pinus brutia, Cupressus sempervirens var. horizontalis, Cupressus arizonica, Pinus eldarica, Elaeagnus angustifolia, Robinia pseudoacacia, Quercus Brantii, Ailanthus altissima, Fraxinus rotundifolia and Pinus nigra in a three hectare forest area in Kamfirooz, Fars province were investigated under RCBD** (Four replicates) for ten years (1993-2003). The area is located at longitude of 52º 10' E and latitude of 32º 22' N. Seedlings were planted at spacing of 3×3m (44 seedling at each) The seedlings survival, collar diameter and height were measured twice a year, at beginning and at end of growth period. The SPSS program, and ANOVA and Duncan tests were used to analyze the data.
The results showed that there were significant differences (p<0.05) between the species performances and there were significant relationship between the species height and diameter and between the species growth parameters and the growth periods. P. brutia and C. arizonica had significantly better growth and survival compared to other species. On the other hand, Pinus nigra showed the least survival, and Q. Brantii showedthe least height and diameter growth.
Overall, it seems that the needleleaved species, except P. nigra,are more successful than thebroadleaved species in respect to their survival, diameter and height parameters.
 
Key words: Diameter, height growth, elimination, species trial, Zagros


Effects of different chemical and livestock fertilizers on growth of Caucasian oak seedlings in two nurseries at Caspian Region of Iran
 
A. Rahmani1 and M. Khoshnevis1
 
Abstract
Two experiments were conducted to determine the effects of N, P and manure fertilizers on growth of Quercus castaneifolia C.A.Meyerin Shahrposht and Koloudeh nurseries (north of Iran). The trial was conducted under factorial statistical method, randomized complete blocks design and three replicates. The factors were Ammonium Sulfate at three levels: 0, 40 and 80 kg/ha), Triple Superphosphate (at three levels: 0, 80 and 120 kg/ha) and manure (at two levels: 0 and 30 T/ha).
The shoot (height and collar diameter) and the root (length, diameter and dry weigh) parameters were measured at each plot, using 15 and tree seedlings, respectively.
In Shahrposht nursery the height and the collar diameter of the seedlings were not affected significantly by any of the fertilizers. Seedlings growth in Koloudeh nursery was improved in fertilized plots, especially with manure application.
The results clearly show that there was significant difference between Koloudeh and Shahrposht nurseries in response to fertilizers.
 
Key words: nursery, fertilization, growth, Caucasian oak, seedling, rootshoot.



1- Member of Scientific Board, Research Center of Agriculture and Natural Resources of Golestan province, Gorgan, I. R. Iran. E- mail: Zaidmirkazemi@ Yahoo.com


2- Member of Scientific Board, Research Institute of Forests and Rangelands, Tehran, I.R. Iran. E-mail: alirezamirbadin@ rifr- ac.org


1- Member of Scientific Board of Azad University. E-mail: moraghebi@yahoo.com


2- Member of Scientific Board of Research Institute of Forests and Rangelands. P.O.Box. 13185-116. Tehran- I.R. Iran.


3- ResearchExperts of Research center of Agricultural and Natural Resources of Gilan province, Gilan, I. R. Iran.


1- Members of scientific Board of Qom Agriculture and Natural Resources Research Center.


1- Member of Scientific Board of Agriculture and Natural Resources Research Center of Markazi Province, E-mail: frahani39@yahoo.com


2- Member of Scientific Board of Research Institute of Forests and Rangelands, Tehran. P. O. Box: 13185-116., Tehran, I. R. Iran.


1- Members of Scientific Board, Fars Research Center of Agriculture and Natural Resources. E-mail: saber@.farsagres.ir


*- Randomized Complete Blocks Design


 


1- Member of scientific board, Research Institute of Forests and Rangelands, Iran. Emaol: arahamani@rifr-ac.ir

کلیدواژه‌ها


عنوان مقاله [English]

A study on morphological, qualitative and quantitative characteristics of oriental beech stands from gene conservation point of view

نویسندگان [English]

  • Parvin Salehi Shanjani 1
  • Khosro Sagheb - Talebi 2
1 ‎- Members of Scientific board, Research Institute of Forests and Rangelands, Tehran, Iran‎
2 Members of Scientific board, Research Institute of Forests and Rangelands,Tehran, Iran.‎
چکیده [English]

Beech is one of the most abundant economically important species in northern Iran. Stand structure identification, combined with silvicultural characteristics, including of ad-mixture, age classes, vertical and horizontal structure, crown cover, absence or presence of gaps and regeneration or under storey could help in optimal management.
Morphological qualitative and quantitative characteristics as well as genetic variation of beech were investigated in 14 sample plots, each covering 0.5 ha; originating from the major part of distribution range of this species in Hyrcanian zone. In this research also the role of human manipulation and selected management methods for management of beech forests on these characteristics were studied.
Considerable genetic diversity of beech forests (expected heterozygosity = 0.191) represents large adaptive potential of this species. Due to positive correlation between genetic diversity and rate of undesirable trees, selection and conservation of desirable trees in shelterwood system must be considered with precautious. Because the selection for conservation of some genotypes is one of the most important processes in forest management practices, it changes the genetic variation of trees. During this process frequency of some genes and following that genetic variability and future adaptability, would be strongly reduced. An appropriate silvicultural system should reflect a sense of conservation, or a determined effort to provide future yields of goods and other values, even while harvesting or using those available at the moment. Therefore, the rate and circumstances of utilizing resources at present would be guaranteed adequate and continuous applies for future. Recent studies revealed that among silvicultural methods, the selection system and group selections are the best options for management of beech forests in a sustainable way. In this research high adaptive potential of some population (Neka at elevation 900 m.a.s.l. and Kheyrud at elevation 600 m.a.s.l. with 14 rare and specific area alleles) in comparison with limited adaptivity of others (such as Kheirud at elevation 1200 m.a.s.l. and Gorgan at elevation 2000 m.a.s.l. with 2 rare and specific area alleles) also emphasized on genetic resource conservation of beech forests and selection of suitable silviculture methods based on potential in every region.
 

کلیدواژه‌ها [English]

  • beech
  • Fagus orientalis Lipsky
  • Iran
  • silviculture
  • Genetic diversity
  • gene conservation
1- اسلامی، ع. ر.، 1378. بررسی ساختار طبیعی راشستانهای خالص و آمیخته در جنگلهای حوزه نکاء ظالمرود. پایان‌نامه کارشناسی ارشد جنگلداری، دانشگاه آزاد اسلامی واحد علوم و تحقیقات، تهران، ایران. 94 صفحه.
2- پارساپژوه، د.، 1355. تحقیق روی کمیتهای فیزیکی چوبهای راش ایران در مناطق مختلف رویشی. مجله منابع طبیعی ایران. 34: 21-34.
3- حبیبی، ح.، 1354. مطالعه وضعیت عناصر (N, P, K, Ca) خاک جنگلهای راش در ایران و بررسی نقش آنها روی رشد راش. مجله منابع طبیعی ایران. 32: 47-62.
4- حبیبی، ح.، 1364. مطالعه خاک جنگلهای راش در ایران و بررسی نقش آن بر توسعه انواع مختلف راش. مجله منابع طبیعی ایران. 39: 6-18.
5- حسینی، س. م. ،1379. تعیین توان اکولوژیکی رویشگاههای سوزنی برگ شمال ایران. رسالة دکتری در دانشگاه تربیت مدرس، تهران، ایران. 162 صفحه.
6- دلفان اباذری، ب.، 1381. بررسی مراحل تحولی و روند پویایی در توده‌های طبیعی دخالت نشده راش در منطقه کلاردشت. رساله دکتری در رشته جنگلداری. واحد علوم و تحقیقات دانشگاه آزاد اسلامی. 210 صفحه.
7- صالحی شانجانی، پ.، 1381. تنوع ژنتیکی راش شرقی و ارتباط آن با برخی ویژگیهای فیزیولوژیکی، بیوشیمیایی و مورفولوژیکی راش در راشستانهای ایران. رساله دکتری دانشکده علوم، دانشگاه تربیت معلم تهران، ایران. 220 صفحه.
8- فلاح، ا.، 1379. بررسی ساختار توده‌های طبیعی راش در استانهای مازندران و گلستان. رساله دکتری در رشته جنگلداری، دانشکده منابع طبیعی، دانشگاه تهران. 200 صفحه.
9- مروی مهاجر، م. ر.، 1355. برخی ویژگیهای کمی جنگلهای راش ایران. مجله منابع طبیعی ایران. 34: 77-97.
10- Crow, J. F. and Kimura, M. 1970. An introduction to population genetics theory. Harper and Row, New York, Evanston, London, 240 p.
11- Eriksson, G., Anderson, S., Eiche, V., Ifver, J. and Persson, A. 1980. Deverity index and transfer effects on survival and volume production of Pinus sylvestris in northern Sweden. Stud. For. Suecica. 156: 1-32.
12- Gøhrn, V., 1972. Proveniens- og afkomsforsøg med bøg (Fagus sylvatica L.). Forstlige Forsgsvaesen Denmark. 33: 83-214.
13- Gregorius, H.R. 1978. The concept of genetic diversity and its formal relationship to heterozygosity and genetic distance. Math Biosci. 41: 253-271.
14- Falkenhagen, E. R. 1985. Isozyme studies in provenance research of forest trees. Theor. Aool. Genet. 69: 335-347.
15- FAO. 2001. International action in the management of forest genetic resources: status and chalenges. Paper compiled by Christel Palmberg-Lerche, march 2001. Forest Genetic Resources Working Papers. Forest resources Development Service, Forest Resources Division. FAO, Rome,
32 p.
16- Hamilton, L. S. 1955. Silvicultural characteristics of American beech. Beech ulitization series, no 13, 25 p.
17- Hamrick, J. L., Godt, M. J. W. and Sherman-Broylers, S. L. 1992. Factors influencing levels of genetic diversity in woody plant species. New Forests. 6: 95-124.
18- Holm, F. 1937. Bøgeracer. Forstlige Forsøgsvaesen Denmark. 14: 193-246.
Kleinschmidt, J. 1977. Forstplanzenzüchtung und Saatgutbereitstellung beim Laubholz. Forst- and Holzwirt. 32: 427-433.
19- Knapp, H. D. and Marvie Mohadjer, R. 2003. The global position of the Caspian forests. International Conference in Mukachevo, Transcarpathia, Ukraine. pp: 82.
20- Krebs, C. J. 1989. Ecology: the experimental analyis of distribution and aboundance. New York, 678 p.
21-Krahl-Urban,1958. Vorläufige Ergebnisse von Buchenprovenien zversuchen. Allg. Forest- und Jagdzeitung. 129: 242-251.
22- Matthewus, J. D. 1989. Silviculture Systems. Clarendon Press. Oxford, 355 p.
23- Merzeau, D., Di Giusto, F., Comps, B., Theibaut, B., and Letouzey, J. and Cuguen, J. 1989. The allozyme variants of beech (Fagus Sylvatica L.): Inheritance and application to a study of the mating system. Silvae Genetica. 38: 195-201.
24- Muona, O. and Szmidt, A. E. 1985. A multilocus study of natural populations of Pinus sylvestris. In Gregorius, H. R. (Eds) population genetics in forestry. Lecture notes in biomathematics. 60: 226-240. Springer-Verlag, Berlin.
25- Müller-Starck, G. and Starke, C. 1993. Auswirkungen von Umweltbelastungen auf genetische Strukturen von Waldbeständen am beispiel der Buche (Fagus sylvatica L.). Schriften aus der Forstl. Fakultät der Universität Göttingen und der Niedersächsischen Forstlichen Versuchsanstalt, J. D. Sauerländers, Frankfurt/M,: 112-193.
26- Nyland, R.D. 1996. Silviculture Concepts and Applications. The mcGraw-Hill,430 p.
27- Peters, R. 1997. Beech forests. Kluwer Academic Publishers. Netherlands. 169 P.
28- Randall, A. 1981. Resource Economics. Wiley, New york , 340 p.
29- Raymond, M. and Rousset, F. 1995b. GENEPOP (Version 1.2): Population genetics software for exact tests and ecumenicism. J. Hered. 86: 248-249.
30- Saghebtalebi, K. 2000. Hyrcanian forests (North of Iran), the unique Ecosystem in Near East region. XXI IUFRO Word Congress-Forests and Society: The Role of research, 7-12 August, 2000, Kuala Lumpur, Malaysia , 141-142.
31- Sagheb-Talebi, K. and Schutz, J. P. 2002. The structure of natural oriental beech (Fagus orientalis) forests in the Caspian region of Iran and potential for the application of the group selection system. Forestry, 75(4): 465  -  472.
32- Smith, M. 1962. The Practice of Silviculture. John Wiley & Sons, Inc., New York, 172 p.
33- Swofford, D. L. and Selender, R. B. 1981. BIOSYS-1: A forstran program for the comprehensive analysis of electrophoretic data in population genetics and systematics. J. Hered. 72: 281-283.
34- Tessier du Cros, E., Kleinschmidt, J., Azoeuf, P. and Hoslin, R. 1980. Spiral grain in beech, variability and heredity. Silva Genetica. 29(1): 5-13.
35- Thiébaut, B., Lumaret, R. and Vernet, P.H. 1982. The bud enzymes of beech (Fagus sylvatica  L..) Genetic distinction and analysis of polymorphism in several French populations. Silvae Genetica, 31, pp. 51 - 60.
36- Tulatrup, N. P. 1950. Proveniensforsøg med nogle vigtige udenlandske bøgeracer. Dansk Skovforenings Tidsskrift. 35: 166 - 178.